Are publication fees the only source of revenue for PLoS?

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Are publication fees the only source of revenue for PLoS?

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In 2003, PLoS embarked on a very ambitious publishing program, and has now established a collection of innovative and high quality open access journals. A new publishing venture like this could never have been launched without substantial financial backing, and we are enormously grateful to our funders for providing the support that we have needed.

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